McVaugh, R. and S. Koptur. 1978. A new species of Pedicularis from Jalisco, Mexico. Contributions to the University of Michigan Herbarium 11: 298-300. [PDF]

Koptur, S. 1979. Facultative mutualism between weedy vetches bearing extrafloral nectaries and weedy ants in California. American Journal of Botany 66(9):
1016-1020. [PDF]

W.A. Haber, G.W. Frankie., H.G. Baker, I. Baker, and S. Koptur. 1981. Ants like flower nectar. Biotropica 13(3): 211-214. [PDF]

Koptur, S., Smith, A.R., and Baker, I. 1982. Nectaries in some neotropical species of Polypodium (Polypodiaceae): preliminary observations and analyses.
Biotropica 14(2): 108-113. [PDF]

Koptur, S. 1983. Flowering phenology and floral biology of Inga (Fabaceae: Mimosoideae). Systematic Botany 8(4): 354-368. [PDF]

Koptur, S. 1983. Inga (Guaba, Guajiniquil, Caite, Paterno). Pp. 259-261 in D.H. Janzen (ed.) Costa Rican Natural History. University of Chicago Press,
Chicago. [PDF]

Koptur, S. 1984. Experimental evidence for defense of Inga (Mimosoideae) saplings by ants. Ecology 65: 1787-1793. [PDF]

Koptur, S. 1984. Outcrossing and pollinator limitation of fruit set: breeding systems of neotropical Inga trees. Evolution 38(5): 1130-1143. [PDF]

Koptur, S. 1985. Alternative defenses against herbivores in Inga (Fabaceae: Mimosoideae) over an elevational gradient. Ecology 66(5): 1639-1650. [PDF]

Law, R. and S. Koptur. 1986. On the evolution of non-specific mutualisms. Biological Journal of the Linnean Society 27: 251-267. [PDF]

Koptur, S. and J.H. Lawton. 1988. Interactions among vetches bearing extrafloral nectaries, their biotic protective agents, and herbivores. Ecology 69: 278-283. [PDF]

Koptur. S. 1988. [Book Review] Ant-Plant Mutualisms and Their Evolution. Evolution 42: 638-639.

Koptur, S., W.A. Haber, G.W. Frankie and H.G. Baker. 1988. Phenological studies of shrub and treelet species in tropical cloud forests of Costa Rica. Journal of
Tropical Ecology 4: 323-346. [PDF]

Koptur, S. 1989. Mimicry of flowers by parasitoid wasp pupae. Biotropica 21(1): 93-95. [PDF]

Koptur, S. 1989. Is extrafloral nectar production an inducible defense? Pp. 323-339 in J. Bock & Y. Linhart (eds.) Evolutionary Ecology of Plants. Westview
Press, Boulder, CO. [PDF]

Koptur, S., E. Davila, D.R. Gordon, B.J. McPhail, C.G. Murphy, and J.B. Slowinski. 1990. The effect of pollen removal on the duration of the staminate phase of
Centropogon talamancensis. Brenesia 33: 15-18. [PDF]

Koptur, S. 1991. Extrafloral nectaries of herbs and trees: modelling the interaction with ants and parasitoids. Pp. 213-229 In D.F. Cutler & C. Huxley (eds.)
Ant-Plant Interactions. Oxford University Press. [PDF]

Koptur, S. 1991. Ecosystems of a Unique and Beleaguered State. [book review of Ecosystems of Florida]. Conservation Biology 5(3): 428-430. [PDF] 

Koptur, S. 1991. Tropical Plant Reproductive Ecology. [book review of Reproductive Ecology of Tropical Forest Plants]. Ecology 72(5): 1911-1912. [PDF]

Koptur, S. 1992. Plants with Extrafloral Nectaries and Ants in Everglades Habitats. The Florida Entomologist 75(1): 38-50. [PDF]

Koptur, S. 1992. Interactions between Insects and Plants Mediated by Extrafloral Nectaries. In Bernays, E. (ed.) CRC series on Insect/Plant Interactions. Volume
4, pp. 85-132. [PDF-first half]
[PDF-second half]

Richards, J.H. and S. Koptur. 1993. Floral variation and distyly in Guettarda scabra (Rubiaceae). American Journal of Botany 80(1): 31-40. [PDF]

Koptur, S. and M.A.B. Lee. 1993. Plantlet formation as a reproductive strategy in tropical ferns. American Fern Journal 83(2): 60-66. [PDF]

Koptur, S. 1994. Floral and extrafloral nectars of neotropical Inga trees: a comparison of their constituents and composition. Biotropica 26(3): 276-284. [PDF]

Koptur, S. 1995. The Everglades - the long and short of it. [book review] Ecology 76(4): 1359-1360.

Oberbauer, S.F., K. Von Kleist III, K.R.T. Whelan, and S. Koptur. 1996. Effects of Hurricane Andrew on epiphyte communities within cypress domes of
Everglades National Park. Ecology 77(3): 964-967. [PDF]

Koptur, S. 1996. The Fascination of Flowers. [book review] Bioscience 46(3): 222-223. [PDF]

Koptur, S. 1996. Extrafloral Nectaries in California Plants: Ecological Implications in Native and Exotic Species. Fremontia 24(2): 23-26. [PDF]

Koptur, S., C.S. Smith, and J.H. Lawton. 1996. Effects of Artificial Defoliation on Reproductive Allocation in the common vetch, Vicia sativa (Fabaceae:  Papilionoideae). American Journal of Botany 83(7): 886-889. [PDF]

Koptur, S. 1997. Floods of Fortune: Ecology and Economy along the Amazon. [book review] The Quarterly Review of Biology 72: 104. [PDF]

Koptur, S., V. Rico-Gray, and M. Palacios-Rios. 1998.  Ant protection in neotropical ferns bearing foliar nectaries.  American Journal of Botany 85(5): 736-739. [PDF]

Koptur, S. and Ni Truong. 1998.  Facultative Ant/Plant Interactions: Nectar sugar preferences of introduced pest ant species in South Florida.  Biotropica 30(2):
179-189. [PDF]

Koptur, S.  1998.  Is a damaged seed a dead seed?  Consequences of seed damage in the common vetch (Vicia sativa L.) and its major herbivore (Cydia spp.).
American Midland Naturalist 140 (2): 393-396. [PDF]

Koptur, S. 1999. [book review] Gardening with a wild heart.  Plant Science Bulletin 45 (3): [PDF]

Koptur, S. 1999.  [book review] Florida Wildflowers in their Natural Communities.  Plant Science Bulletin 45 (3): [PDF]

 Koptur, S. 2000.  Breeding systems of Monteverde Inga.  Pp. 85-87 in Nadkarni, N.M. and N.T. Wheelwright (eds.) Monteverde:  ecology and conservation of a tropical cloud forest.  Oxford University Press, New York. [PDF]

Koptur, S. 2000.  Interactions among Inga, herbivores, ants, and insect visitors to foliar nectaries.  Pp. 277-278 in Nadkarni, N.M. and N.T. Wheelwright (eds.) Monteverde:  ecology and conservation of a tropical cloud forest.  Oxford University Press, New York. [PDF]

 Koptur, S. 2000. [book review] Britain’s Rare Flowers.  Plant Science Bulletin 46(4):132. [PDF]

 Koptur, S. 2001. [book review] Warming up to Hotspots.  Conservation Biology 15 (2): 542-545.

Koptur, Suzanne.  2002.  “Old Herbaceous” - H. G. Baker - A warm and light heart.  Tropinet 13 (2):  2-3. [weblink]

Koptur, S., M. C. Rodriguez, S. F. Oberbauer, C. Weekley, and A. Herndon.  2002. Herbivore-Free Time?  Damage to New Leaves of Woody Plants After Hurricane Andrew.  Biotropica 34(4): 547-554. [PDF]

 Liu, H. and S. Koptur. 2003.  Breeding System and Pollination of a Narrowly Endemic Herb of the Lower Florida Keys: Impacts of the Urban Wildland Interface.  American Journal of Botany  90: 1180-1187. [PDF]

 Sah, J.P., M. S. Ross, S. Koptur and J. R. Snyder.  2004.   Estimating aboveground biomass of broadleaved woody plants in the understory of Florida Keys pine forests.  Forest Ecology and Management 203: 319-329. [PDF]

Liu, H., E. S. Menges, J. R. Snyder, S. Koptur, M. S. Ross.  2005.  Effects of fire intensity on vital rates of an endemic herb of the Florida Keys, USA.  Natural Areas Journal 25 (1): 71 – 76. [PDF]

Koptur, S.  2005. Nectar as fuel for plant protectors.  Ch. 3 (pp. 75-108) in F.L. Wäckers, P.C.J. van Rijn & J. Bruin (editors) Plant-provided food for carnivorous insects:  a protective mutualism and its applications.  Cambridge University Press. [PDF]

Cairns, C. E., R. Villanueva-Gutiérrez, S. Koptur, and D. B. Bray.  2005. Bee populations, forest disturbance, and Africanization in Mexico.  Biotropica 37 (4): 686-692.  [PDF]

Koptur, S.  2006. The conservation of specialized and generalized pollination systems in subtropical ecosystems:  a case study.  Ch. 15 in N. Waser and J. Ollerton (eds.) Plant–pollinator interactions: from specialization to generalization.  University of Chicago Press. [PDF].

Koptur, S. 2006.  H.G. Baker.  (Biographical Memoirs).  Proceedings of the American Philosophical Society 150 (2): 335-339. [PDF]

Sah, J.P., M.S. Ross, J.R. Snyder, S. Koptur, and H.C. Cooley. 2006.  Fuel loads, fire regimes, and post-fire fuel dynamics in Florida Keys pine forests.  International Journal of Wildland Fire 15: 1-16. [PDF]
Koptur, S. 2007.  The Everglades Handbook [book review of 2nd edition, 2005, by Thomas E. Lodge].  The Palmetto 24 (3): 15. [PDF]

Koptur, S. 2007.  Natural Enemies: an Introduction to Biological Control. [book review of 2004 book by Ann Hajek]. Plant Science Bulletin 53(2):73.  [PDF]

Pemberton, R., T. Collins, and S. Koptur. 2008.  An Asian orchid, Eulophia graminea (Orchidaceae:  Cymbidieae), naturalizes in Florida.  Lankesteriana 8(1): 5-14. [PDF]

Pemberton, R., S. Koptur, and T. Collins.  2008.  A Newly Naturalized Orchid found in Florida.  P. 412-413 in Orchids, June 2008.  [PDF]  

Pemberton, R., S. Koptur, and T. Collins. 2008.  Alien terrestrial orchid, Eulophia graminea, invades Miami.  North American Native Orchid Journal 14 (3): 183-184. [PDF]

Khorsand Rosa, R., and S. Koptur. 2009.  Preliminary observations and analyses of pollination in Coccothrinax argentata:  do insects play a role?  Palms 53 (2): 75-83. [

Elena Pinto-Torres and Suzanne Koptur. 2009.  Hanging by a coastal strand: breeding system of a federally endangered morning-glory of the south-eastern Florida coast, Jacquemontia reclinata.  Annals of Botany 104 (7): 1301-1311 [PDF]; doi: 10.1093/aob/mcp241  available from publisher's link  

Koptur, S. 2009.  A Zapotec Natural History – Trees, Herbs, and Flowers, Birds, Beasts, and Bugs in the Life of San Juan Gbëë. [book review of 2008 book by Eugene S. Hunn]. Plant Science Bulletin 55(2): 81-2. [PDF

Koptur, S.  2009.  Seedling Ecology and Evolution. [book review of 2008 book edited by Mary Leck, Parker, and Simpson]. Plant Science Bulletin 55(4): 169 – 170.  [PDF]
Amalin, D.M., J.E. Peña, R. Duncan, J. Leavengood, and S. Koptur. 2009.  Effects of pesticides on the arthropod community in the agricultural areas near the Everglades National Park.   Proc. Fla. State Hort. Soc. 122:429–437. 2009. [PDF

Cardel, Y. and S. Koptur. 2010. Effects of florivory on the pollination of flowers: an experimental field study with a perennial plant.  International Journal of Plant Science 171(3):283–292. [PDF

Koptur, S., P. William, and Z. Olive. 2010.  Ants and plants with extrafloral nectaries in fire successional habitats on Andros (Bahamas).  Florida Entomologist 93(1): 89-99. [PDF

Koptur, S. 2010.  Seedlings of Barro Colorado Island and the tropics [book review of 2009 book by Nancy Garwood].  Plant Science Bulletin 56(1): 36-7. [PDF

Gamper, H.A. and S. Koptur. 2010. Honeydew foraging by birds in tropical montane forests and pastures of Mexico.  Journal of Tropical Ecology 26:335–341. [PDF

Linares, L.J. and S. Koptur. 2010. Floral biology and breeding system of the crenulate lead plant, Amorpha herbacea var. crenulata, an endangered south Florida pine rockland endemic.  Natural Areas Journal 30:138-147. [

Geiger, J.H., P.D. Pratt, and S. Koptur. 2010.  Breeding system of Ruellia succulenta Small (Acanthaceae).  Castanea 75(4): 454-460. [

Barrios, B. and S. Koptur. 2011.  Floral biology and breeding system of Angadenia berteroi (Apocynaceae):  why do flowers of the pineland golden trumpet produce few fruits?  International Journal of Plant Sciences 172(3): 378-385. [

Barrios, B., G. Arellano, and S. Koptur. 2011. The effects of fire and fragmentation on occurrence and flowering of a rare perennial plant.  Plant Ecology 212: 1057-1067. [

Koptur, S. 2011. A long, productive, and inspiring career [book review of Jean Langenheim's autobiography entitled The Odyssey of a Woman Field Scientist: a story of passion, persistence, and patience].  Ecology 92 (5): 1190-91. [PDF]

Gamper HA, Koptur S, Garcia-Franco J, Stapper AP. 2011.  Alteration of forest structure modifies the distribution of scale insect, Stigmacoccus garmilleri, in Mexican tropical montane cloud forests.  Journal of Insect Science 11: 120 (14 pp.). available online:  [PDF]

 Maschinski, Joyce, Samuel J. Wright , Suzanne Koptur, and Elena C. Pinto-Torres. 2013. Is local the best paradigm? Breeding history influences conservation reintroduction survival and population trajectories in times of extreme climate events. Biological Conservation 159: 277-284. [PDF]

Khorsand Rosa, Roxaneh, and Suzanne Koptur. 2013. New findings on the pollination biology of Mauritia flexuosa (Arecaceae) in Roraima, Brazil: linking dioecy, wind, and habitat.  American Journal of Botany 100 (3): 613–621. [PDF]

Suzanne Koptur, Mónica Palacios-Rios, Cecilia Díaz-Castelazo, William P. Mackay, and Víctor Rico-Gray. 2013. Nectar secretion on fern fronds associated with lower levels of herbivore damage: field experiments with a widespread epiphyte of Mexican cloud forest remnants. Annals of Botany 111(6): 1277-1283 doi:10.1093/aob/mct063. [PDF]

 Roxaneh Khorsand Rosa, Reinaldo Imbrozio Barbosa, and Suzanne Koptur. 2013. How do habitat and climate variation affect phenology of the Amazonian palm, Mauritia flexuosa?  Journal of Tropical Ecology, 29, pp 255-259. doi:10.1017/S0266467413000242. [PDF]

 Brigitte Marazzi, Judith L. Bronstein, and Suzanne Koptur. 2013. The diversity, ecology and evolution of extrafloral nectaries: current perspectives and future challenges. Annals of Botany 111(6): 1243-1250. doi:10.1093/aob/mct109. [PDF]

 Koptur S., J.E. Peña, and W. L. Grogan, Jr.  2013. The biting midge, Forcipomyia (Microhelea) eriophora (Williston) (Diptera: Ceratopogonidae), an ectoparasite of larval Phoebis sennae (Pieridae) in south Florida.  Journal of the Lepidopterists’ Society 67(2): 128–130.  [PDF]  

Jones, Ian M. and S. Koptur. 2015. Dynamic extrafloral nectar production: the timing of leaf damage affects the defensive response in Senna mexicana var. chapmanii Fabaceae). American Journal of Botany 102(1): 58-66. [PDF]

Dewsbury, Bryan M., Suzanne Koptur, and James W. Fourqurean. 2015. Ecosystem responses to prescribed fire along a chronosequence in a subtropical pine rockland habitat. Caribbean Naturalist 24: 1-12. [PDF]

Jones, Ian M. and S. Koptur. 2015. Quantity over quality: light intensity, but not red/far-red ratio, affects extrafloral nectar production in Senna mexicana var. chapmanii. Ecology and Evolution (open access) doi: 10.1002/ece3.1644 [PDF]

Koptur, Suzanne, Ian M. Jones, and Jorge E. Peña. 2015. The influence of host plant extrafloral nectaries on multitrophic interactions: an experimental investigation.  PLoS ONE 10(9): e0138157. doi:10.1371/journal.pone.0138157 [PDF]

Soti, Pushpa, K. Jayachandran, S. Koptur, and J. Volin. 2015. Effect of soil pH on growth, nutrient uptake, and mycorrhizal colonization in exotic invasive Lygodium microphyllum. Plant Ecology 05/2015; 216(7). DOI:10.1007/s11258-015-0484-6  [PDF]

Barrios, Beyte, Sean R. Pena, Andrea Salas, and Suzanne Koptur. 2016.  Butterflies visit more frequently, but bees are better pollinators: the importance of mouthpart dimensions in effective pollen removal and deposition. AoB PLANTS 8: plw001; doi:10.1093/aobpla/plw001 [PDF]

 Beyte Barrios Roque, Suzanne Koptur, and Jay P. Sah. 2016. The effects of habitat fragmentation on the reproduction and abundance of Angadenia berteroi. Journal of Plant Ecology 10 (2): 340-348. [PDF]

Jones, I. M., Koptur, S. and von Wettberg, E. J. 2017. The use of extrafloral nectar in pest management: overcoming context dependence. Journal of Applied Ecology 54: 489–499. doi:10.1111/1365-2664.12778 [PDF]

Jones, I. M., S. Koptur, H. R. Gallegos, J. P. Tardanico, P. A. Trainer, and J. Peña. 2017. Changing light conditions in pine rockland habitats affect the intensity and outcome of ant/plant interactions. Biotropica 49:83-91. [PDF]

Clayborn, Jaeson, Suzanne Koptur, George O’Brien, and Kevin R. T. Whelan. 2017. The Schaus Swallowtail Habitat Enhancement Project: An Applied Service-Learning Project Continuum from Biscayne National Park to Miami-Dade County Public Schools. Southeastern Naturalist (SENA) 16 (Outdoor Classroom Special Issue 10):26–46. [PDF]

Suzanne Koptur and Alexandra Dutton. 2017. The FIU Nature Preserve: Achieving Biological Conservation Through Diversification. Southeastern Naturalist (SENA) 16 (Outdoor Classroom Special Issue 10):94–108. [PDF]

Jones, I., S. Koptur, and J.E. Peña. 2017. Exploring whether and how ants affect reproductive fitness in Senna mexicana var. chapmanii. Florida Entomologist 100(3): 539-545. [PDF]

Koptur, S. and D. Garcia. 2017. Habitat Differences in Morphology and Reproductive Allocation in Guettarda scabra (Rubiaceae). Castanea 82(2): 51–57. [PDF]

 Clayborn, Jaeson, and Suzanne Koptur. 2017. Mortal combat between ants and caterpillars: An ominous threat to the endangered Schaus swallowtail butterfly (Heraclides aristodemus ponceanus) in the Florida Keys, USA. Journal of Insect Conservation 21(4): 689-702. DOI: 10.1007/s10841-017-0012-1. [PDF]

Koptur, Suzanne. 2017. Plant-Animal Interactions in the Face of Disturbance (Keynote Address). Pp. 15-25, 1st Joint Symposium on the Natural History and Geology of the Bahamas, edited by Lee Florea, Carol Landry, Daniel Kjar. Gerace Research Center, San Salvador, Bahamas. [PDF]

Oliveira, P.S. and S. Koptur (editors). 2017. Ant-Plant Interactions: Impacts of Humans on Terrestrial Ecosystems. Cambridge University Press. 456 pp. [PDF]

Suzanne Koptur, Ian M. Jones, Hong Liu, and Cecilia Diaz-Castelazo. 2017. Playing the system: the impacts of invasive ants and plants on facultative ant-plant interactions.   Chapter 12 in Paulo Oliveira and Suzanne Koptur (eds.) Ant-Plant Interactions - Impacts of Humans on Terrestrial Ecosystems. Cambridge University Press.   [PDF]

Jones, Ian M. and Suzanne Koptur. 2018. Dead land walking: the value of continued conservation efforts in South Florida’s imperiled pine rocklands. Biodiversity and Conservation ISSN 0960-3115 Biodivers Conserv DOI 10.1007/s10531-017-1433-6. [PDF]

 Koptur, S. and R. Khorsand. 2018. Pollination Ecology of Three Sympatric Palms of Southern Florida Pine Rocklands. Natural Areas Journal 38: 15-25. [PDF]

Koptur, S., T. Livshultz, G. Ionta, and C-W Tan. 2019. Contrasting Pollination Systems of Wild Unction and Devil's Potato Root (Apocynaceae) on San Salvador: Preliminary Observations and Analyses. Castanea 84(1): 81-92. URL:  [PDF]          

Gonsiska, P. and S. Koptur. 2019. Mystery herbivore of charismatic epiphyte discovered on Andros. Journal of the Lepidopterist’s Society 73(2), 2019, xx–xx. [PDF]

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Other Publications by graduate students on research done under my supervision:

O’Brien, J.J. 1998.  The distribution and habitat preferences of rare Galactia species (Fabaceae) and Chamaesyce deltoidea subspecies (Euphorbiaceae) native to southern Florida pine rockland.  Natural Areas Journal 18 (3): 208-222.

Smith, E.M. 2002.  The effects of season, host plant protection, and ant predators on the survival of Eumaeus atala (Lycaenidae) in re-establishments.  J. Lepidopterist’s Society 56(4): 272-276.

Spira, L. P. and J.R. Snyder. 1998.  Effects of wet- and dry-season fires on Jacquemontia curtisii, a south Florida pine forest endemic.  Natural Areas Journal 18 (4): 350-357.

Weekley, Jr., C.W. 2000.  The natural history of Tanaostigmodes pithecellobiae (Hymenoptera:  Tanaostigmatidae), a gall-maker on blackbead (Pithecellobium keyense).  Florida Entomologist 83 (1): 31-41

Khorsand Rosa, R. 2011. Bees collect resin from Mauritia flexuosa in Roraima, Brazil. Palms 55(4): 200-203.

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Popular articles or mentions of our work in the popular press:

Biology by the Month. Monteverde Mountain Monthly. October 1979 [PDF]

Ants, plants, and conniving caterpillars.  BBC Wildlife April 1988 pp. 192-3. [PDF]

Moths, butterflies are prominent pollinators - many bugs are transportation for propagation. Georgia Tasker, Miami Herald, Sunday 29 September 1996. [PDF]

The key: Stop thinking of insects as pests. Georgia Tasker, Miami Herald, Sunday 29 September 1996. [PDF]

Nature triumphs against the odds. Cyril T. Zaneski, Miami Herald, Monday 18 August 1997. [PDF]

What good can nectar do a fern?  1998. Science News 153: 344. [PDF]

FNPS 23rd Annual Conference Highlights. Palmetto 22(3):9. [PDF]

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